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55 it a better proxy to avoid detection bias. Moreover, body pits are considered the most accurate proxy to describe patterns in reproductive output after the direct count of individuals (SWOT Scientific Advisory Board 2011). Tableau 3. Designation and geographic positions of the 28 surveyed sites in the GLS rookery. The geographic coordinates point the center of the islet. islet # name latitude longitude set islet # name latitude longitude set 1 atire -22.5219 166.5603 A 15 nge -22.6926 166.8507 A 2 redika -22.5161 166.6130 A 16 gi -22.7212 166.8516 A 3 vua -22.5781 166.7475 A 17 nda -22.8456 166.8773 A 4 uo -22.5169 166.7696 B 18 uie -22.7208 166.9125 B 5 mato -22.5503 166.7923 B 19 mbore -22.8030 166.9161 B 6 uatio -22.7090 166.7953 A 20 ugo -22.4417 166.9285 B 7 kouare -22.7767 166.8009 A 21 koko -22.8804 166.9329 B 8 ua -22.7079 166.8112 A 22 petit koko -22.8584 166.9439 B 9 uaterembi -22.6795 166.8117 A 23 ndo -22.6840 166.9653 B 10 noe -22.5393 166.8218 B 24 totea -22.7203 166.9735 B 11 ieroue -22.6049 166.8277 A 25 nouare -22.4238 167.0128 B 12 puemba -22.5321 166.8289 B 26 kie -22.3761 167.0646 B 13 pumbo -22.5205 166.8355 B 27 amere -22.4441 167.0961 B 14 tere -22.7888 166.8475 A 28 du ami -22.6252 167.2797 B 3.2.2 Monitoring strategy in space and time The ‘GLS’ rookery presents a spatial configuration where numerous, non-contiguous, distant and hard-to-reach nesting beaches are presumably used by the same nesting population. Setting a well-adapted monitoring scheme is challenging, since no preconized protocol reflects the entire complexity of the present situation (SWOT Scientific Advisory Board 2011). Given the logistic impossibility to monitor each islet with high intensity throughout the nesting season, the initial selection of 28 islets has been divided into two distinct sets, respectively designated as A- and B-set, that will differ in the total seasonal number of survey occurrences (Figure 16 ; Tableau 3).

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